* the incorporation of new developments in phylogenetics, developmental biology, and molecular genetics
* major changes at the highest levels among the invertebrates. Three phyla that appeared in the original book—Pentastomida, Pogonophora, and Vestimentifera—no longer exist, and a new phylum, Cycliophora, has been erected. Moreover, this edition discusses recent work in molecular systematics that has shaken classic views on animal classification.
* a large new section on "Kingdom Protista" (replacing "Protozoa") containing new contemporary views of these organisms (arranged in 18 phyla).
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